| Hummingbird Temporal range: Rupelian to present | |
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| Four hummingbirds from Trinidad and Tobago | |
Hummingbirds are birds from the Americas that constitute the family Trochilidae. They are among the smallest of birds, most species measuring 7.5–13 cm (3–5 in) in length. Indeed, the smallest extant bird species is a hummingbird, the 5 cm (2.0 in) bee hummingbird weighing less than 2.0 g (0.07 oz).
They are known as hummingbirds because of the humming sound created by their beating wings which flap at high frequencies audible to humans. They hover in mid-air at rapid wing-flapping rates, which vary from around 12 beats per second in the largest species, to in excess of 80 in some of the smallest. Of those species that have been measured in wind tunnels, their top speed exceeds 15 m/s (54 km/h; 34 mph) and some species can dive at speeds in excess of 22 m/s.
Hummingbirds have the highest metabolism of any homeothermic animal. To conserve energy when food is scarce, and nightly when not foraging, they can go into torpor, a state similar to hibernation, slowing metabolic rate to 1/15th of its normal rate.
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Evolution
A map of the hummingbird family tree—reconstructed from analysis of 284 of the world's 338 known species—shows rapid diversification from 22 million years ago. Hummingbirds fall into nine main clades, the Topazes, Hermits, Mangoes, Brilliants, Coquettes, Patagona, Mountain Gems, Bees, and Emeralds, defining their relationship to nectar-bearing flowering plants and the birds' continued spread into new geographic areas.
While all hummingbirds depend on flower nectar to fuel their high metabolisms and hovering flight, coordinated changes in flower- and bill shape stimulated the formation of new species of hummingbirds and plants. Due to this exceptional evolutionarypattern, as many as 140 hummingbird species can coexist in a specific region, such as the Andes range.
The hummingbird evolutionary tree shows ancestral hummingbirds splitting from insectivorous swifts (family Apodidae) and tree swifts (family Hemiprocnidae) about 42 million years ago, probably in Eurasia. One key evolutionary factor appears to be an altered taste receptor that enabled hummingbirds to seek nectar. By 22 million years ago the ancestral species of current hummingbirds became established in South America, where environmental conditions stimulated further diversification.
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Vision
During evolution, hummingbirds have adapted to the navigational needs of visual processing while in rapid flight or hovering by development of an exceptionally dense array of retinal neurons allowing for increased spatial resolution in the lateral and frontal visual fields. Morphological studies showed that neuronal hypertrophy, relatively the largest in any bird, exists in a brain region called the pretectal nucleus lentiformis mesencephali (or nucleus of the optic tract in mammals) responsible for refining dynamic visual processing while hovering and during rapid flight. The enlargement of this brain region responsible for visual processing indicates enhanced ability for perception and processing of fast-moving visual stimuli which hummingbirds encounter during rapid forward flight, insect foraging, competitive interactions, and high-speed courtship.
Hummingbirds are highly sensitive to stimuli in their visual fields, responding to even minimal motion in any direction by reorienting themselves in midflight. Hummingbirds' visual sensitivity allows them to precisely hover in place while in complex and dynamic natural environments, functions enabled by the lentiform nucleus which is tuned to fast-pattern velocities, enabling highly tuned control and collision avoidance during forward flight.
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Flight stability
During turbulent airflow conditions created experimentally in a wind tunnel, hummingbirds exhibit stable head positions and orientation when they hover at a feeder. When wind gusts from the side, hummingbirds compensate by increasing the amplitude of their wing strokes plane angle and by varying the orientation and enlarging the collective surface area of their tail feathers into the shape of a fan. While hovering, the visual system of a hummingbird is able to separate apparent motion caused by the movement of the hummingbird itself from motions caused by external sources, such as an approaching predator. In natural settings full of highly complex background motion, hummingbirds are able to precisely hover in place by rapid coordination of vision with body position.
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Feather colors
To serve courtship and territorial competition, many male hummingbirds have plumage with bright, varied coloration resulting both from pigmentation in the feathers and from prism-like cells within the top layers of feathers of the head, gorget, breast, back and wings. When sunlight hits these cells, it is split into wavelengths that reflect to the observer in varying degrees of intensity, with the feather structure acting as a diffraction grating. Iridescen the hummingbird colors result from a combination of refraction and pigmentation, since the diffraction structures themselves are made of melanin, a pigment, and may also be colored by carotenoid pigmentation and more subdued black, brown or gray colors dependent on melanin.
By merely shifting position, feather regions of a muted-looking bird can instantly become fiery red or vivid green. In courtship displays for one example, males of the colorful Anna's hummingbird orient their bodies and feathers toward the sun to enhance the display value of iridescent plumage toward a female of interest.
One study of Anna's hummingbirds found that dietary protein was an influential factor in feather color, as birds receiving more protein grew significantly more colorful crown feathers than those fed a low-protein diet.Additionally, birds on a high-protein diet grew yellower (higher hue) green tail feathers than birds on a low-protein diet
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Tongue as a micropump
Hummingbirds drink with their tongues by rapidly lapping nectar. Their tongues have tubes which run down their lengths and help the hummingbirds drink the nectar. While capillary action was believed to be what drew nectar into these tubes, high-speed photography has revealed that the tubes open down their sides as the tongue goes into the nectar, and then close around the nectar, trapping it so it can be pulled back into the beak. The tongue, which is forked, is compressed until it reaches nectar, then the tongue springs open, the rapid action traps the nectar and the nectar moves up the grooves, like a pump action, with capillary action not involved. Consequently, tongue flexibility enables accessing, transporting and unloading nectar.
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In myth and culture[edit]
- Aztecs wore hummingbird talismans, both artistic representations of hummingbirds and fetishes made from actual hummingbird parts: emblematic for their vigor, energy, and propensity to do work along with their sharp beaks that symbolically mimic instruments of weaponry, bloodletting, penetration, and intimacy. Hummingbird talismans were prized as drawing sexual potency, energy, vigor, and skill at arms and warfare to the wearer.[
- The Aztec god of war Huitzilopochtli is often depicted as a hummingbird (right). It was also believed that fallen warriors would return to earth as hummingbirds and butterflies. The Nahuatl word huitzil (hummingbird) is an onomatopoeic word derived from the sounds of the hummingbird's wing-beats and zooming flight.
- One of the Nazca Lines depicts a hummingbird (right).
- Trinidad and Tobago, known as "The land of the hummingbird," displays a hummingbird on that nation's coat of arms, 1-cent coin[132] and emblem of its national airline, Caribbean Airlines...
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